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Lars O. Hedin is the George M. Moffett Professor of Biology in the Department of Ecology and Evolutionary Biology at Princeton University. Research in his laboratory centers on ecosystem analysis, with emphasis on the emergence and maintenance of geographically broad patterns in cycling of nutrients and greenhouse trace gases. Postdoctoral, graduate, and undergraduate students work on diverse topics that range from local-scale microbial processes to global-scale controls on ecosystem structure and function. Current interests fall in three general areas: 1) Broad controls on nutrient cycles in temperate and tropical forests; 2) Emergence of macroscopic properties (e.g., stoichiometric ratios and ecosystem functions) from Darwinian selection; and 3) Biophysical controls on soil-atmosphere exchange of the greenhouse gas methane.